https://hdl.handle.net/10481/68932 2026-04-21T18:38:42Z 2026-04-21T18:38:42Z Otero, José Carlos https://hdl.handle.net/10481/80029 2023-02-17T10:29:25Z

The world fauna of the genus Micrambe Thomson, 1863 (Coleoptera, Cryptophagidae) Otero, José Carlos The species of the genus Micrambe Thomson, 1863 (Coleoptera, Cryptophagidae) are revised. Hundred and one species are recognized. Key and figures of the studied species are provided. Following a detailed study of the type material in different entomological collections, the status of taxa is revised and 3 new synonyms are proposed: Micrambe (Micrambinus) schuelkei Esser, 2017a, Micrambe (Micrambinus) yunnanensis Esser, 2017 and Micrambe (Micrambinus) zhejiangensis Esser, 2017 are nov. syn. of Micrambe (Micrambinus) bimaculata (Panzer, 1798). Micrambe duclouxi Grouvelle, 1910 and Micrambe duclouxi (Grouvelle, 1916) are new as synonymy of Micrambe (Micrambe) sinensis Grouvelle, 1910; Micrambe aubrooki Donisthorpe, 1939 is new synonymy with Micrambe hirta Grouvelle, 1908. Micrambe (Micrambe) sinensis Grouvelle, 1910 is resurrected from synonymy with Micrambe (Micrambinus) bimaculata (Panzer, 1798). Micrambe (Micrambe) micramboides (Reitter, 1874) previously known only from Japan is reported here for the first time from Taiwan.; Se revisan las especies del género Micrambe Thomson, 1863 (Coleoptera, Cryptophagidae). Se recogen ciento una especies. Se proporcionan claves y figuras de las diferentes especies estudiadas. Tras un estudio detallado del material tipo en diferentes colecciones entomológicas, se revisa el estado de los taxones y se proponen tres nuevos sinónimos: Micrambe (Micrambinus) schuelkei Esser, 2017a, Micrambe (Micrambinus) yunnanensis Esser, 2017 y Micrambe (Micrambinus) zhejiangensis Esser, 2017 para Micrambe (Micrambinus) bimaculata (Panzer, 1798). Micrambe duclouxi Grouvelle, 1910 y Micrambe duclouxi (Grouvelle, 1916) son ahora sinónimas de Micrambe (Micrambe) sinensis Grouvelle, 1910; Micrambe aubrooki Donisthorpe, 1939 es ahora sinónima de Micrambe hirta Grouvelle, 1908. Micrambe (Micrambe) sinensis Grouvelle, 1910 es rescatada de su sinonimia con Micrambe (Micrambinus) bimaculata (Panzer, 1798). Además, Micrambe (Micrambe) micramboides (Reitter, 1874), previamente citada en Japón, se reporta por primera vez de Taiwán.

Villastrigo, Adrián Millán, Andrés Sánchez Fernández, David Fresneda, Javier Valladares, Luis F. https://hdl.handle.net/10481/78007 2022-12-07T23:23:17Z

Advances in aquatic and subterranean beetles research: a tribute to Ignacio Ribera Villastrigo, Adrián; Millán, Andrés; Sánchez Fernández, David; Fresneda, Javier; Valladares, Luis F. It has been a bit longer than two years since our friend and colleague Ignacio (Nacho) Ribera passed away. The memory of him remains among those of us who were lucky enough to meet Nacho. This monograph is dedicated to him, bringing a set of scientific contributions from his colleagues whose topics are part of the main research lines (and passions) of his scientific work: taxonomy, systematics, biogeography and evolution of aquatic and subterranean beetles. In the last two years, several contributions have highlighted Nacho’ scientific and personal profile, including the complete list of his publications, and both the taxa described by and dedicated to him (BELLÉS, 2020; CEHRE, 2020; DELOCADO et al., 2020; FAILLE et al., 2020, 2021; JÄCH, 2020; MELIC, 2020; MILLÁN et al., 2020a, b; VALLADARES & MILLÁN, 2020). Nacho was a passionate biologist, interested in everything concerning beetles, especially their diversity, distributions, adaptations and evolution of aquatic and cave beetles. He published a total of 285 papers, in which he described 107 species new to science, highlighting the discovery of the Aspyditidae family. Besides, 7 species and subspecies, and one new genus have been dedicated to him in this monograph, extending to a total of 15 species and two genera. Concerning the aquatic beetles, we would like to emphasise his crucial contributions, such as the first complete checklist of aquatic and semi-aquatic beetles of the Iberian Peninsula. He was a pioneer in studying the adaptive morphology of the family Dytiscidae, and more importantly, he deepens in the phylogeny of the aquatic beetles, shedding light on the systematics and evolution of the families Dytiscidae and Hydraenidae. He postulated the “Habitat Constraint” hypothesis which, from an evolutionary point of view, highlights the importance of habitat stability as a determinant of species range sizes (via differences in their dispersal capabilities). He also dealt in-depth with evolutionary studies related to the habitat transition between lotic and lentic water bodies, but also the transition from freshwater to saline water, or vice versa. Certainly, Nacho, due to his work and publications on aquatic beetles, became one of the most prestigious specialists worldwide. Regarding the subterranean environment, Nacho revealed that Dalyat Mateu is a vicariant genus of carabid whose origin must be in the separation of the Iberian plate from the rest of Pangea in the Jurassic-Cretaceous boundary. He also addressed the position of Ildobates neboti Español, confirming it within the Zuphiini tribe. At the same time, he began to study the phylogeny of the hypogeal Trechini of the Pyrenees, and afterwards he addressed the diversification of Troglocharinus Reitter, opening the way to a new perspective on the evolution and dispersion of the subterranean fauna. He published the first morphological phylogeny through cladistic analysis of the Leptodirini tribe (Leiodidae), and the first molecular phylogenies for the two main groups that have colonised the subterranean environment, the tribes Leptodirini and Trechini. His contribution to the knowledge of underground and stygobic environments between 2005 and 2021, postulated him as one of the most important specialists worldwide also in this field. The complete list of scientific works signed by Nacho, the taxa described by him and those dedicated to his person can be found on the web: https://www.um.es/ecoaqua/index.php/external-collaborators. We do not want to extend further. We hope these pages serve as a tribute to his person, but also a tribute to a life and scientific style that would serve as an example for future generations of insect lovers. Our gratitude to the authors and reviewers of the articles that make up this monograph, as well as to the Asociación española de Entomología, which has greatly facilitated its preparation, edition and publication.

Vela, José Miguel Daccordi, Mauro https://hdl.handle.net/10481/72692 2022-02-07T10:14:37Z

A study of the genus Timarcha Samouelle, 1819: the species from the South of the Iberian Peninsula (Coleoptera, Chrysomelidae) Vela, José Miguel; Daccordi, Mauro The species of Timarcha Samouelle, 1819 living in the South of the Iberian Peninsula are reviewed basing on the available types. We have recognized twelve species-group taxa: Timarcha apricaria ssp. apricaria Waltl, 1835, T. apricaria ssp. heydeni Weise, 1882, T. apricaria ssp. parvicollis Rosenhauer, 1856, T. granadensis Bechynĕ, 1948, T. insparsa Rosenhauer, 1856, T. intermedia ssp. intermedia Herrich-Schäffer, 1838, T. intermedia ssp. lugens Rosenhauer, 1856, T. kiesenwetteri ssp. kiesenwetteri Kraatz, 1879, T. kiesenwetteri ssp. sagrensis Kuntzen, 1911, T. marginicollis Rosenhauer, 1856, T. scutellaris Waltl, 1835 and T. strophium ssp. carmelenae Petitpierre, 2013. Other two taxa studied here, Timarcha apricaria ssp. melitensis Weise, 1882 and T. strophium ssp. strophium Weise, 1888, are spread all around North Morocco (Rif) and Central-Eastern Spain (Cuenca and Teruel) respectively. The identity of the long-time unknown T. apricaria and T. scutellaris has been re-established. Timarcha heydeni, T. parvicollis and T. melitensis have been downgraded to a subspecific rank of T. apricaria, and T. carmelenae has been downgraded to a subspecific rank of T. strophium. Timarcha lugens is confirmed as a subspecies of T. intermedia, and T. kiesenwetteri ssp. sagrensis is tentatively maintained as a subspecies until more specimens will be available. The following new synonyms are proposed: Timarchaapricaria ssp. apricaria = T. coarcticollis Fairmaire & Allard, 1873, nov. syn. = T. paulinoi Kraatz, 1879, nov. syn. = T. parvicollis ssp. nitentula Bechynĕ, 1953, nov. syn. Timarcha apricaria ssp. parvicollis = T. seidlitzi Kraatz, 1879, nov. syn. = T. parvicollis ssp. elocata Bechynĕ, 1953, nov. syn. Timarcha apricaria ssp. melitensis = T. riffensis Fairmaire, 1888, nov. syn. Timarcha intermedia ssp. lugens = T. parnassia Weise, 1882 nov. syn. = T. nevadensis Fairmaire, 1884, nov. syn. Timarcha scutellaris = T. hispanica Herrich-Schäffer, 1838, nov. syn. = T. rugipennis Perez Arcas, 1865, nov. syn. = T. erosa Fairmaire & Allard, 1873, nov. syn. = T. vermiculata Fairmaire, 1880, nov. syn. = T. laevisterna Fairmaire, 1880, nov. syn. = T. transversicollis Fairmaire 1884, nov. syn. Timarcha strophium ssp. strophium = T. fossulata Bechynĕ, 1944, nov. syn. Timarcha splendida Pérez-Arcas, 1872 and T. sericea var. escalerae Kuntzen, 1911, are confirmed as synonyms of T. marginicollis Rosenhauer, 1856. Timarcha strophium Weise, 1888 has been redescribed basing on the types. The type series of Timarcha granadensis ssp. dislocata Bechynĕ, 1948 is made up by two species: one of them is T. granadensis; the other is a species from Central-Eastern Spain close to T. sobrina Fairmaire, 1884. Neotypes have been designated for Timarcha hispanica, T. intermedia and T. scutellaris. Lectotypes have been designated for Timarcha apricaria, T. erosa, T. granadensis, T. heydeni, T. insparsa, T. kiesenwetteri, T. laevisterna, T. lugens, T. melitensis, T. nevadensis, T. parnassia, T. parvicollis ssp. elocata, T. parvicollis ssp. nitentula, T. paulinoi, T. riffensis, T. rugipennis, T. seidlitzi, T. sericea, T. splendida, T. strophium and T. vermiculata. Timarcha apricaria Gistel, 1832 and T. angulicollis Motschulsky, 1849 are considered nomina dubia, and two species described as native of Andalusia, T. gravis Rosenhauer, 1856 and T. rugosula Rosenhauer, 1856, are alien to the Iberian fauna

Serrano, José Andújar, Antonio Lencina, José Luis https://hdl.handle.net/10481/68939 2021-06-15T18:27:40Z

Taxonomy and biogeography of Iberian Amarina Bonelli, 1810 (Coleoptera, Carabidae) Serrano, José; Andújar, Antonio; Lencina, José Luis The systematics of Iberian Amarina Bonelli, 1810 (Coleoptera, Carabidae) is revised by updating the catalogue of species and their distribution. Keys to determine the genera, subgenera and species are included. To date, 73 species and subspecies included in 13 subgenera and 2 genera are known from Iberia. It is discussed the position of Curtonotus Stephens, 1828 and Amarocelia Motschulsky, 1862 as separate taxa in the light of new evidence. The biogeography of Amarina is also discussed; species of this subtribe show a wide distribution within the Palearctic region, whereas the whole family Carabidae shows a higher proportion of endemisms and Mediterranean elements in the Iberian Peninsula. Comments on the evolutionary history of Amarina are likewise included; the group seems to have an Asiatic European origin and colonized the Mediterranean basin when the ancient Tethys Sea was closed.; Se ha revisado la sistemática de los Amarina Bonelli, 1810 ibéricos (Coleoptera, Carabidae), actualizando el catálogo de especies y su distribución y se ha elaborado claves para la determinación de géneros, subgéneros y especies. Hasta la fecha se conocen en la península Ibérica 73 especies y subespecies de Amarina, que se incluyen en 13 subgéneros y 2 géneros. Se aportan argumentos recientes y clásicos para considerar a Curtonotus Stephens, 1828 y Amarocelia Motschulsky, 1862 como taxones independientes. También se discuten aspectos biogeográficos de los Amarina ibéricos; el patrón de los taxones de esta subtribu es diferente al del conjunto de la familia Carabidae en la península ibérica, ya que la mayoría de los Amarina presentan una distribución amplia en la región Paleártica, mientras que en el conjunto de los carábidos ibéricos predominan los endemismos y los elementos mediterráneos. También se incluyen consideraciones sobre la historia evolutiva de toda la subtribu Amarina, que parece ser un grupo asiático europeo cuya llegada a la cuenca mediterránea se produjo al cerrase el antiguo mar de Tethys. Thanks are due to colleagues of the Department of Zoology of Murcia for providing beetles included in this study, and the personnel of the Museo Nacional de Ciencias Naturales (Madrid) in charge of beetle collections (particularly Carolina Martín), for the facilities to consult hundreds of specimens. We also thank our colleagues Juan and Eduard Vives, Marcos Toribio and Juan Pérez Zaballos for donating valuable specimens included in this study. Thanks are also due to Francisco Rodríguez Luque (Faluke) and Fran García for providing the colour photographs of Amara species. The study has been supported by the project Animal Phylogeny and Evolution (project no. 19908/GERM/15) funded by the Fundación Séneca, Murcia. We have no competing interests in the elaboration of this paper.

Verves, Yuri G. https://hdl.handle.net/10481/68937 2021-06-15T18:27:39Z

An annotated list of Chinese Sarcophagidae (Insecta, Diptera) Verves, Yuri G. An annotated list of 105 genera and 343 species, including 315 species from People’s Republic of China (267 species from Palaearctic and 127 from Oriental parts), 63 species from Taiwan and 34 species from Tibet is presented. 9 new combinations of species names are published: Cylindrothecum brachygonitum (Fan & Ge, 1992), comb. nov.; C.curticlaws (Zhang & Liu, 1998), comb. nov.; Miltogrammidium fidusum (Wei & Yang, 2007), comb. nov.; M. indigenum (Wei & Yang, 2007), comb. nov.; M. leigongshanum (Wei & Yang, 2007), comb. nov.; Asceloctella (s. str.) australis (Johnston & Tiegs, 1921), comb. nov.; Sarcorohdendorfia pudongensis (Fan, Chen & Lu, 2003), comb. nov.; S. shenzhenfensis (Fan, 2002), comb. nov and Fanzidea kurahashii (Shinonaga & Tumrasvin, 1979), comb. nov. 17 new synonyms are established: Kozlovea lopesi Nandi, 1976 = Kozlovea yangiana Lehrer, 2010, syn. nov.; K. tshernovi Rohdendorf, 1937 = Kozlovea nyatria Lehrer, 2010, syn. nov.; Leucomyia alba (Schiner, 1868) = Leucomyia tarima Lehrer, 2008, syn. nov.; Harpagophalla kempi (Senior-White, 1924) = Harpagophalla samudera Lehrer, 2008, syn. nov.; H. kempi (Senior-White, 1924) = Harpagophalla papuasia Lehrer, 2008, syn. nov.; H. kempi (Senior-White, 1924) = Harpagophalla nandiana Lehrer, 2008, syn. nov.; Thyrsocnema (Pseudothyrsocnema) ugamskii Rohdendorf, 1937 = Asiopierretia amuriella Lehrer, 2009, syn. nov.; T. (P.) ugamskii Rohdendorf, 1937 = Asiopierretia henania Lehrer, 2011, syn. nov.; Myorhina villeneuvei (Böttcher, 1912) = Myorhina altainula Lehrer, 2009, syn. nov.; Pseudothyrsocnema caudagalli (Böttcher, 1912) = Pierretia autochtona Wei et Yang, 2007, syn. nov.; Fengia ostindicae (Senior- White, 1924) = Fengia shanga Lehrer, 2008, syn. nov.; Liosarcophaga (Jantiella) javana (Macquart, 1851) = Jantiella bentenia Lehrer, 2008, syn. nov.; Robineauella (s. str.) nigribasicosta (Ye, 1982) = Sarcophaga (Robineauella) picibasicosta Pape, 1996, syn. nov.; Sarcorohdendorfia antilope (Böttcher, 1913) = Tricholioproctia wumengia Lehrer et Wei, 2011, syn. nov.; S. inextricata (Walker, 1860) = Tricholioproctia sulawesiella Lehrer et Wei, 2011, syn. nov.; Seniorwhitea princeps (Wiedemann, 1830) = Seniorwhitea fuhsia Lehrer, 2008, syn. nov. and Phallosphaera gravelyi (Senior-White, 1924) = Phallosphaera yelangiops Lehrer et Wei, 2011, syn. nov.; Se presenta una lista comentada de 105 géneros y 343 especies, de las cuales 315 especies son de la República Popular China (267 del paleártico y 127 de la región oriental), 63 especies de Taiwán y 34 especies del Tíbet. Se publican nueve nuevas combinaciones de nombres de especies: Cylindrothecum brachygonitum (Fan & Ge, 1992), comb. nov.; C.curticlaws (Zhang & Liu, 1998), comb. nov.; Miltogrammidium fidusum (Wei & Yang, 2007), comb. nov.; M. indigenum (Wei & Yang, 2007), comb. nov.; M. leigongshanum (Wei & Yang, 2007), comb. nov.; Asceloctella (s. str.) australis (Johnston & Tiegs, 1921), comb. nov.; Sarcorohdendorfia pudongensis (Fan, Chen & Lu, 2003), comb. nov.; S. shenzhenfensis (Fan, 2002), comb. nov and Fanzidea kurahashii (Shinonaga & Tumrasvin, 1979), comb. nov.. Se establecen 17 nuevas sinonimias: Kozlovea lopesi Nandi, 1976 = Kozlovea yangiana Lehrer, 2010, syn. nov.; K. tshernovi Rohdendorf, 1937 = Kozlovea nyatria Lehrer, 2010, syn. nov.; Leucomyia alba (Schiner, 1868) = Leucomyia tarima Lehrer, 2008, syn. nov.; Harpagophalla kempi (Senior-White, 1924) = Harpagophalla samudera Lehrer, 2008, syn. nov.; H. kempi (Senior-White, 1924) = Harpagophalla papuasia Lehrer, 2008, syn. nov.; H. kempi (Senior-White, 1924) = Harpagophalla nandiana Lehrer, 2008, syn. nov.; Thyrsocnema (Pseudothyrsocnema) ugamskii Rohdendorf, 1937 = Asiopierretia amuriella Lehrer, 2009, syn. nov.; T. (P.) ugamskii Rohdendorf, 1937 = Asiopierretia henania Lehrer, 2011, syn. nov.; Myorhina villeneuvei (Böttcher, 1912) = Myorhina altainula Lehrer, 2009, syn. nov.; Pseudothyrsocnema caudagalli (Böttcher, 1912) = Pierretia autochtona Wei et Yang, 2007, syn. nov.; Fengia ostindicae (Senior-White, 1924) = Fengia shanga Lehrer, 2008, syn. nov.; Liosarcophaga (Jantiella) javana (Macquart, 1851) = Jantiella bentenia Lehrer, 2008, syn. nov.; Robineauella (s. str.) nigribasicosta (Ye, 1982) = Sarcophaga (Robineauella) picibasicosta Pape, 1996, syn. nov.; Sarcorohdendorfia antilope (Böttcher, 1913) = Tricholioproctia wumengia Lehrer et Wei, 2011, syn. nov.; S. inextricata (Walker, 1860) = Tricholioproctia sulawesiella Lehrer et Wei, 2011, syn. nov.; Seniorwhitea princeps (Wiedemann, 1830) = Seniorwhitea fuhsia Lehrer, 2008, syn. nov. y Phallosphaera gravelyi (Senior-White, 1924) = Phallosphaera yelangiops Lehrer et Wei, 2011, syn. nov. DISCUSSION The results of studying of sarcophagids of People’s Republic of China, Tibet and Taiwan are presented in Appendix 1. The differences in sizes and natural conditions of different administrative unites are unreliable for constatation of real degree of studiing of their faunas. Total number of sarcophagids’ species is 343, i. e. for People’s Republic of China – 315, for Tibet – 34 and Taiwan – 63. Only Taiwan fauna is known detaily; other two faunas have been studied not more than about of 80-90%. The number of known species from Palaearctic part of China (267) more than twice exceeded as such from Oriental part (128). The largest numbers of species are known from Neimenggu (92), Sichuan (87), Liaoning (84), Xinjiang (75), Yunnan (73), Shanxi (61), Jilin (58), and Guangdong (53). Those datae compose from 25 to 46 species for majority of provinces, and below 20 only for 4 ones: Jiangxi (17), Hong Kong (16), Anhui (8), and Tianjin (6).; I wish to express my sincere thanks to my friend and colleague, Dr. Sci., Prof. Wanqi XUE (Institute of Entomology, Shenyang Normal University, Shenyang, China) for sending of the literature necessary to compile the present catalogue and standing collaboration, and to all reviewers for detailed text check review and numerous corrections of text.